Russia's women's 4x400metres relay squad has been stripped of a London 2012 Olympic silver medal after Antonina Krivoshapka tested positive, the International Olympic Committee has announced.

The IOC on Wednesday announced three further anti-doping rule violations following re-analysis of samples from Beijing 2008 and London 2012, taking the total to 101.

Krivoshapka tested positive for the steroid turinabol and her sixth-placed finish in the 400m individual event and Russia's second-placed finish in the relay event have been wiped from the record books.

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The United States won the relay event and Jamaica finished third, with Ukraine fourth.

Russian discus thrower Vera Ganeeva tested positive for the same substance and her 23rd-placed finish at London 2012 has been expunged from the records.

Turkish boxer Adem Kilicci, who reached the quarter-finals of the middleweight (67-75kg), also tested positive for turinabol and was disqualified.

The three positive tests took the total to 40 adverse findings from 492 samples re-analysed from the London 2012 Olympics.

Of the 1,053 samples re-analysed from the Beijing 2008 Olympics, 61 have been positive to date.